Access Denied to Address Book Folder

After installing a new Lync 2013 Standard Edition server, the event log continually presented the following error:
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Address Book Server has encountered an unexpected exception.

Exception: Access to the path '\\LyncServerFQDN\LyncFileShare\1-WebServices-2\ABFiles\00000000-0000-0000-0000-000000000000\00000000-0000-0000-0000-000000000000' is denied.

Exception Type: System.UnauthorizedAccessException

   at System.IO.__Error.WinIOError(Int32 errorCode, String maybeFullPath)
   at System.IO.Directory.InternalCreateDirectory(String fullPath, String path, Object dirSecurityObj, Boolean checkHost)
   at System.IO.Directory.InternalCreateDirectoryHelper(String path, Boolean checkHost)
   at Microsoft.Rtc.ABServer.ABServer.ReadContactsFromSqlAndWriteToOutput(NormalizeRuleSet normalizationRules, SqlConnection sqlConnection, List`1 sinks)
   at Microsoft.Rtc.ABServer.ABServer.Synchronize()
Cause: Internal Error
Resolution:

Contact Product Support Services.

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To resolve the issue I had to manually add the following local system groups to the folders sharing permissions:

LOCAL\RTC Local Administrators                  CHANGE,READ

LOCAL\RTC Local Config Replicator             CHANGE,READ

LOCAL\RTC Server Local Group                    CHANGE,READ

LOCAL\RTC Component Local Group            CHANGE,READ

After adding these and running Update-CsUserDatabase I immediately could see the creation of the expected Address Book files.

Checking other deployments these permissions were not listed, so I am unsure why in this case they were required.